1. Milleret, C., Dupont, P., Dey S., Brøseth H., Kindberg J., Turek D., de Valpine P., Åkesson M., Wabakken P., Zimmermann B. and Bischof R. (2025) Map of death: spatially explicit mortality of the grey wolf. Proceedings of the Royal Society. B.29220250948

2. Sanz-Pérez, A., Sazatornil, V., Palazón, S., Vanpé, C., Quenette, P. Y., Sentilles, J., Kervellec, M., Milleret, C., … & Sollmann, R. (2025). Mapping sex-and age-structure reveals lonely males at the front in an expanding brown bear population. Biological Conservation, 306, 111122.

3. Moqanaki, E., Milleret, C., Dupont, P., Mattisson, J., Dey, S., Brøseth, H., … & Bischof, R. (2025). Environmental variability across space and time drives the recolonization pattern of a historically persecuted large carnivore. Proceedings of the National Academy of Sciences, 122(5), e2401679122.

4. Ausilio, G., Sand, H., Wikenros, C., Aronsson, M., Milleret, C, Nordli, K., … & Zimmermann, B. (2024). Effects of large carnivores, hunter harvest, and weather on the mortality of moose calves in a partially migratory population. Wildlife Biology, 2024(6), e01179.

5. Milleret, C., Dupont, P., Chapron, G., Swenson, J. E., & Bischof, R. (2024). Sweden is shooting brown bears in the dark. Science, 385(6713), 1052-1052.

6. Boiani, M. V., Dupont, P., Bischof, R., Milleret, C., Friard, O., Geary, M., Avanzinelli, E., von Hardenberg, A., & Marucco, F. (2024). When enough is enough: Optimising monitoring effort for large-scale wolf population size estimation in the Italian Alps. Ecology and Evolution, 14, e70204. https://doi.org/10.1002/ece3.70204

7. Marucco, F., Boiani, M. V., Dupont, P., Milleret, C., Avanzinelli, E., Pilgrim, K., … & Bischof, R. (2023). A multidisciplinary approach to estimating wolf population size for long‐term conservation Conservation Biology, 37(6), e14132. , e01179.

8. Edelhoff, H., Milleret, C., Ebert, C., Dupont, P., Kudernatsch, T., Zollner, A., … & Peters, W. (2023) Sexual segregation results in pronounced sex-specific density gradients in the mountain ungulate, Rupicapra rupicapra Communications Biology, 6(1), 979. , e01179.

9. Moqanaki, E., Milleret, C., Dupont, P., Brøseth, H., & Bischof, R. (2023). Wolverine density distribution reflects past persecution and current management in Scandinavia Ecography , (87) e06689.

10. Neby, M., Andreassen, H., Milleret, C., P., Pedersen, S., Tamayo, A. M. P., Sánchez, D. C., … & Zimmermann, B. (2023). Small rodent monitoring at Birkebeiner road, Norway. Biospanersity Data Journal, 11, e105914.

11. Kervellec, M., Milleret, C., Vanpé, C., Quenette, P. Y., Sentilles, J., Palazón, S., … & Gimenez, O. (2023). An evaluation of spatial capture-recapture models applied to ungulate non-invasive genetic sampling data Biological Conservation , (87) e22373.

12. Dupont, P., Bischof, R., Milleret, C., Peters, W., Edelhoff, H., Ebert, C., Klamm, A., and Hohmann, U. (2023) An evaluation of spatial capture-recapture models applied to ungulate non-invasive genetic sampling data Journal of Wildlife Management , (87) e22373.

13. Tourani, M., Franke, F., Heurich, M., Henrich, M., Peterka, T., Ebert, C., Oeser, J., Edelhoff, H., Milleret, C., Dupont, P., Bischof, R., Peters, W., (2023) Spatial variation in red deer density in a transboundary forest ecosystem, Scientific Reports , (13) 4513.

14. Dey, S., Moqanaki, E., Milleret, C., Dupont, P., Tourani, M., Bischof, R., (2023) Modelling spatially autocorrelated detection probabilities in spatial capture-recapture using random effects Ecological Modelling (479) 110324.

15. Milleret, C., Dey, S., Dupont, P., Brøseth, H., Turek, D., de Valpine P., Bischof, R., (2022) Estimating spatially variable and density-dependent survival using open-population spatial capture–recapture models Ecology 104(2): e3934

16. Zhang, W., Chipperfield, J., Illian, J., Dupont, P., Milleret, C., De Valpine, p., Bischof, R, (2022) A flexible and efficient Bayesian implementation of point process models for spatial capture‐recapture data Ecology 104(1): e3887

17. Semper-Pascual, A., Bischof, R., Milleret, C., et al. (2022) Occupancy winners in tropical protected forests: a pantropical analysis Proceedings of the royal society B 2892022045720220457

18. Theng, M., Milleret, C., Bracis, C., Cassey, P., Delean, S. (2022) Confronting spatial capture–recapture models with realistic animal movement simulations Ecology 12(7):e3676

19. Soumen, D., Dupont, P., Bischof, R., Milleret, C., (2022) Does the punishment fit the crime? Consequences and diagnosis of misspecified detection functions in Bayesian spatial capture–recapture modeling Ecology and Evolution 12(2):e8600

20. Dupont, P., Milleret, C., Tourani, M., Brøseth, H., Bischof, R. (2021) Integrating dead recoveries in open-population spatial capture–recapture models Ecosphere 12(7):e03571

21. Milleret, C., Bischof, R., Dupont, P., Brøseth, H., Odden, J., and Mattison, J. (2021) GPS collars have an apparent positive effect on the survival of a large carnivore Biol. Lett 17(20210128)

22. Moqanaki, E., Milleret, C., Tourani, M., Dupont, P., Bischof, R. (2021) Consequences of ignoring variable and spatially autocorrelated detection probability in spatial capture-recapture Landscape Ecology 12(2):e03385

23. Turek, D., Milleret, C., Ergon, T., Brøseth, H., Dupont, P., Bischof, R., de Valpine P., (2021) Efficient estimation of large‐scale spatial capture–recapture models Ecosphere 12(2):e03385

24. Crampe, JP., Gerard, JF., Goulard, M., Milleret, C., Gonzalez, G., Bon, R. (2021) Year-round sexual segregation in the Pyrenean chamois, a nearly monomorphic polygynous herbivore Behavioural Processes 184, 104300

25. Bischof, R., Milleret, C., Dupont, P., Chipperfield, J., Tourani, M., Ordiz, A., de Valpine, P., Turek, D., Royle, A., Gimenez, O., Flastad, Ø., Åkesson, M., Svensson, L., Brøseth, H., Kindberg, J. (2020) Estimating and forecasting spatial population dynamics of apex predators using transnational genetic monitoring Proceedings of the National Academy of Sciences

26. Ordiz, A., Milleret, C., Uzal, A., Zimmermann, B., Wabakken, P., Wikenros, C., Sand, H., Swenson, J., Kindberg, J. (2020) individual variation in predatory behavior, scavenging and seasonal prey availability as potential drivers of coexistence between wolves and bears spanersity. 12, 356.

27. Milleret, C., Dupont, P., Chipperfield, J., Turek, D., Brøseth, H., Gimenez, O., de Valpine, P., Bischof, R. (2020) Estimating abundance with interruptions in data collection using open population spatial capture–recapture models Ecosphere. 11(6):e03172.

28. Ordiz, A., Uzal, A., Milleret, C., Sanz-Pérez A., Zimmermann, B., Wikenros, C., Wabakken, P., Kindberg, J., Swenson, J., Sand, H. (2020) Wolf habitat selection when sympatric or allopatric with brown bears in Scandinavia Scientific Reports. 10, 9941.

29. Carricondo-Sanchez, D., Zimmermann, B., Wabakken, P., Eriksen, A., Milleret, C., Ordiz, A., Sanz-Pérez, A., Wikenros K. (2020) Wolves at the door? Factors influencing the individual behavior of wolves in relation to anthropogenic features Biological Conservation. 9:8849.

30. Bischof, R., Dupont, P., Milleret, C., Chipperfield, J., Royle J.A. (2020) Consequences of ignoring group association in spatial capture–recapture analysis. Wildlife Biology. (1).

31. Sanz‐Pérez, A., Giralt, D., Robleño, I., Bota, G., Milleret, C., Mañosa, S., & Sardà‐Palomera, F. (2019). Fallow management increases habitat suitability for endangered steppe bird species through changes in vegetation structure. Journal of Applied Ecology, 56(9), 2166-2175.

32. Bischof, R., Gjevestad, J. G. O., Ordiz, A., Eldegard, K., & Milleret, C. (2019). High frequency GPS bursts and path-level analysis reveal linear feature tracking by red foxes. Scientific reports, 9(1), 8849.

33. Donfrancesco, V., Ciucci, P., Salvatori, V., Benson, D., Andersen, L. W., Bassi, E., Milleret, C, … & Mukherjee, N. (2019). Unravelling the scientific debate on how to address wolf-dog hybridization in Europe. Frontiers in Ecology and Evolution, 7, 175.

34. Dupont, P., , Gimenez, O., & Bischof, R. (2019). Population closure and the bias‐precision trade‐off in spatial capture–recapture. Methods in Ecology and Evolution, 10(5), 661-672.

35. Milleret, C., Ordiz, A., Sanz-Perez, A., Uzal, A., Carricondo-Sanchez, D., Eriksen, A., … & Zimmermann, B. (2019). Testing the influence of habitat experienced during the natal phase on habitat selection later in life in Scandinavian wolves. Scientific Reports, 9(1), 6526.

36. Milleret, C., Dupont, P., Bonenfant, C., Brøseth, H., Flagstad, Ø., Sutherland, C., & Bischof, R. (2019). A local evaluation of the individual state‐space to scale up Bayesian spatial capture–recapture. Ecology and evolution, 9(1), 352-363.

37. Sanz-Pérez, A., Ordiz, A., Sand, H., Swenson, J. E., Wabakken, P., Wikenros, C., … & Milleret, C. (2018). No place like home? A test of the natal habitat-biased dispersal hypothesis in Scandinavian wolves. Royal Society Open Science, 5(12), 181379.

38. Milleret, C., Ordiz, A., Chapron, G., Andreassen, H. P., Kindberg, J., Månsson, J., … & Sand, H. (2018). Habitat segregation between brown bears and gray wolves in a human‐dominated landscape. Ecology and Evolution, 8(23), 11450-11466.

39. Milleret, C., Dupont, P., Brøseth, H., Kindberg, J., Royle, J. A., & Bischof, R. (2018). Using partial aggregation in spatial capture recapture. Methods in Ecology and Evolution, 9(8), 1896-1907.

40. Tallian, A., Ordiz, A., Metz, M. C., Milleret, C., Wikenros, C., Smith, D. W., … & Sand, H. (2017). Competition between apex predators? Brown bears decrease wolf kill rate on two continents. Proceedings of the Royal Society B: Biological Sciences, 284(1848), 20162368.

41. Milleret, C., Wabakken, P., Liberg, O., Åkesson, M., Flagstad, Ø., Andreassen, H. P., & Sand, H. (2017). Let’s stay together? Intrinsic and extrinsic factors involved in pair bond dissolution in a recolonizing wolf population. Journal of Animal Ecology, 86(1), 43-54.

42. Chapron, G., Wikenros, C., Liberg, O., Wabakken, P., Flagstad, Ø., Milleret, C, … & Sand, H. (2016). Estimating wolf (Canis lupus) population size from number of packs and an individual based model. Ecological Modelling, 339, 33-44.

43. Ordiz, A., Milleret, C, J. Kindberg, J. Månsson, P. Wabakken, J. E. Swenson, and H. Sand. 2015. Wolves, people, and brown bears influence the expansion of the recolonizing wolf population in Scandinavia. Ecosphere 6(12):284. http://dx.doi.org/10.1890/ES15-00243.1

44. Nicholson, K.L., Milleret, C., Månsson, J. et al. (2014) Testing the risk of predation hypothesis: the influence of recolonizing wolves on habitat use by moose. Oecologia 176, 69–80.

Technical reports

1. Duchamp, C,. Milleret, C., Gimenez, O., Augeard, B., Séon-Massin, N (2025). Etat de conservation du loup en France : mise à jour 2025 de la viabilité démographique de la population sous régime de tirs dérogatoires - Office Français de la Biodiversité. 2025, 25 p. ⟨hal-05274113

2. Dupont, P., Milleret, C., Semper-Pascual, A., Brøseth, H., Kopatz, A., Kindberg, J. and Bischof R. (2025). Estimates of brown bear density, abundance, and population dynamics in Norway 2015 - 2024 - MINA fagrapport 105. 35pp.

3. Semper-Pascual, A., Dupont, P., Milleret, C., and Bischof, R., (2025). Spatial variation in detectability of bears, wolverines and wolves during large-scale non-invasive genetic sampling in Norway and Sweden- MINA fagrapport 104. 27 pp.

4. Svensson, L., Wabakken, P., Maartmann, E., Nordli, K., Jensen, M. Ø., Milleret, C, … & Flagstad, Ø. (2025). Bestandsovervåking av ulv vinteren 2024-2025.

5. Milleret, C., Dupont, P., Semper-Pascual, A., Brøseth, H., Flagstad, Ø., Kindberg, J., Svensson, L., and Bischof, R., (2025). Estimates of wolf density, abundance, and population dynamics in Sweden and Norway, 2015–2025- MINA fagrapport 103. 36 pp.

6. Milleret, C., Duchamp, C., Gimenez, O., Mise à jour des estimations démographiques et des effectifs de la population de loups en France lors de l’hiver 2023/2024. (2025). CNRS; OFB. .hal-05016361

7. Milleret, C., Dupont, P., Brøseth, H., Flagstad, Ø, Kleven, O., Königsson, H., Spong, G., Kindberg, J., and Bischof, R., 2024. Estimates of wolverine density, abundance, and population dynamics in Scandinavia, 2015–2024- MINA fagrapport 101. 34 pp.

8. Milleret, C., Dupont, P., Winiger, A., Spong, G., Königsson, H., and Bischof, R., (2024). Estimates of wolverine density and abundance in Norrbotten county in Sweden and associated reindeer herding areas, 2023/2024 - MINA fagrapport 100. 20 pp.

9. Dupont, P., Milleret, C., Brøseth, H., Kindberg, J. and Bischof, R. (2024). Estimates of brown bear density, abundance, and population dynamics in Norway 2014 - 2023 - MINA fagrapport 98. 34pp.

10. Milleret, C., Dupont, P., Brøseth, H., Flagstad, Ø., Kindberg, J., Svensson, L., and Bischof, R., (2024).Estimates of wolf density, abundance, and population dynamics in Scandinavia, 2014–2024- MINA fagrapport 97. 34 pp.

11. Milleret, C., Dupont, P., Åsbrink, J., and Bischof, R., (2024). Simultaneous range-wide genetic sampling of brown bears in Sweden: a pilot study - MINA fagrapport 96. 23 pp.

12. Dupont, P., Milleret, C., Brøseth, H., Kindberg, J., and Bischof, R., (2024). Challenges in estimating range-wide brown bear density and abundance in Sweden (2012-2021) - MINA fagrapport 95. 47pp.

13. Milleret, C., Dupont, P., Brøseth, H., Flagstad, Ø., Kleven, O., Kindberg, J., & Bischof, R. (2023). Estimates of wolverine density, abundance, and population dynamics in Scandinavia, 2014–2023. MINA fagrapport, 89, 33 pp.

14. Milleret, C., Dupont, P., Brøseth, H., Flagstad, Ø., Kindberg, J., Svensson, L., & Bischof, R., (2023). Estimates of wolf density, abundance, and population dynamics in Scandinavia, 2013–2023. MINA fagrapport, 85, 34 pp.

15. Dupont, P., Milleret, C., Brøseth, H., Kindberg, J., & Bischof, R. (2023). Estimates of brown bear density, abundance, and population dynamics in Norway 2012 - 2022. MINA fagrapport, 86, 33pp.

16. Dupont, P., Milleret, C., Brøseth, H., Kindberg, J., & Bischof, R. (2022) Estimates of brown bear density, abundance, and population dynamics in Norway 2012-2021 - MINA fagrapport, 82, 32pp.

17. Milleret, C., Dupont, P., Åkesson, M., Svensson, L., Brøseth, H., Kindberg, J., & Bischof, R. (2022). Estimates of wolf density, abundance, and population dynamics in Scandinavia, 2013–2022. MINA fagrapport, 77, 35pp.

18. Milleret, C., Dupont, P., Moqanaki, E., Brøseth, H., Flagstad, O., Kleven, O., … & Bischof, R. (2022). Estimates of wolverine density, abundance, and population dynamics in Scandinavia, 2014–2022. MINA fagrapport, 79, 35pp.

19. Milleret, C., Dupont, P., Brøseth, H., Flagstad, O., Kindberg, J., & Bischof, R. (2022). Estimates of wolverine density, abundance, and population dynamics in Scandinavia, 2013–2021. MINA fagrapport, 74, 30pp.

20. Milleret, C., Dupont, P., Åkesson, M., Brøseth, H., Kindberg, J., & Bischof, R. (2021). Estimates of wolf density, abundance, and population dynamics in Scandinavia, 2012-2021. MINA fagrapport 72. 30pp

21. Milleret, C., Dupont, P., Åkesson, M., Svensson, L., Brøseth, H., & Bischof, R. (2020). Consequences of reduced sampling intensity for estimating population size of wolves in Scandinavia with spatial capture-recapture models. MINA fagrapport 65. 29pp

22. Bischof, R., Milleret, C., Dupont, P., Chipperfield, J., Brøseth, H., & Kindberg, J. (2019). RovQuant: Estimating density, abundance and population dynamics of bears, wolverines and wolves in Scandinavia.MINA fagrapport 63. 79pp

23. Bischof, R., Milleret, C., Dupont, P., Chipperfield, J., Åkesson, M., Brøseth, H., & Kindberg, J. (2019). Estimating the size of the Scandinavian wolf population with spatial capture-recapture and conversion factors. MINA fagrapport 57. 80 pp

24. Chapron, G., Liberg, O., Wikenros, C., Svensson, L., Åkesson, M., Månsson, J., Zimmermann, B., Milleret, C, Sand, H. (2015). Population estimates for the Scandinavian wolf population and sample-based monitoring. Rapport från Viltskadecenter, (2015: 5).